The U.S. military said Thursday it had captured the leaders of a Shiite insurgent network responsible for one of the boldest and most sophisticated attacks on American troops since the Iraq conflict began four years ago.

The statement said the arrests took place over the past three days in the cities of Basra and Hillah south of Baghdad. The military said the network was led by Qais Khazaali and his brother Laith Khazaali. Several other members of the network also were captured.

The network was "directly connected" to the killing in January of five American soldiers in the holy city of Karbala, 50 miles (80 kilometers) south of Baghdad, the military said.

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