Yes, if one wanted to develop notions of "cultural evolution" these would be things that would have to be addressed -- replicators, unit individuals, etc. But with respect to features that promote adaptability, one can black-box the question of replicators and say, whatever form they may take, here are the systemic traits needed to promote their ability to adapt. I think there may be some clear sailing along that line of thought (famous last words...).
Bob
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